Notes - Notes : SMTP Protocol Returned a Permanent Error 554 Relay rejected for policy reasons

We have a BES server which works fine for incoming and outgoing mail to Blackberry handhelds OTHER than when a user does a Repy to an email sent from the internal Notes domain.

When they reply to an internal Notes user the mail fails in the mail.box on the BES server with an error “SMTP Protocol Returned a Permanent Error 554 Relay rejected for policy reasons”. However, the mail recipient is structured as:

Joe Blogss/DomainName@DomainName

Any idea why this is failing?